小魔仙夏乐彤、颜可可、白尹玥带着她们的伙伴穿梭精灵布丁、咪噜和鹿比开启了蕴藏着四季能量的灵春之庭、幻夏之屿和暖秋之苑。她们以守护者的身份，维护着四季的平衡。来自凛冬之崖的守护者司齐因受到蒙蔽，意图冰封另外三个季节，恢复凛冬秩序，小魔仙一边守护着四季，一边经历成长中的各种烦恼，在此过程中，她们学会接受自己，包容他人，更全面的看待问题，同时也加深了彼此之间的羁绊，最终用默契和勇气反败为胜，化解了寒冬危机。

Mary Howard (Crawford) is a novelist with advanced ideas about love and marriage, and is in love with her publisher, Rogers Woodruff (Marshall). She decides the only logical thing to do is to lure him away from his wife and marry him. Mary's friend, Jimmy (Taylor), however, is convinced he's the right man for her and pursues her. He sees through her rationalizations and wrong-thinking and decides to throw Mary and Woodruff's wife Claire (Garson) together at the house of a friend (Byington). The two women do not know each other, but during their chats Mary appreciates and respects Claire's maturity and wisdom. When she learns Woodruff is a philandering womanizer of long standing, she realizes she cannot love him and welcomes Jimmy's attentions.

Mary, a writer working on a novel about a love triangle, is attracted to her publisher. Her suitor Jimmy is determined to break them up; he introduces Mary to the publisher's wife without telling Mary who she is.

小魔仙夏乐彤、颜可可、白尹玥带着她们的穿梭精灵伙伴，布丁、咪噜和鹿比，守护着灵春之庭、幻夏之屿和暖秋之苑。来自凛冬之崖的守护者司齐，在夜落的蒙蔽和蛊惑下，帮助他突破了冰墙封印。恢复自由的夜落为了获取强大的自然能量，抢夺蝶梦星，企图让黑暗笼罩四季。悔恨不已的司齐为了对抗夜落，加入小魔仙，并进入星缘中学学习知识。开阔了眼界的他，逐渐发现自然的奥妙，汇聚了属于自己的蝶梦星。在这个过程中，他和小魔仙彼此包容，共同对抗夜落，建立了新的羁绊。四季循环，生生不息，最终，她们凭借强大的意志，和守护四季的信念，集齐十二颗蝶梦星，用四季的自然能量，打败夜落，化解了黑暗危机。

The difficulty of establishing beauty and the futility of power.

秦灭汉起，百废待兴。诸子隐世，氏族势行。驱车逐鹿，弱肉何争？同心携力，主臣兼行。天命赐异，源于甘星，施于阴阳，异于静动。静者天行，缓而止争；动者袍泽，速若雷霆。捕星之计，谋策之心。甘星入世，衍造异人。奸者祸国，忠者祸身，道貌离合，莫再成臣。离经叛道，天下重分。采薇采薇，复始如周，采薇采薇，复始若秦。

秦灭汉起，百废待兴。诸子隐世，氏族势行。驱车逐鹿，弱肉何争？同心携力，主臣兼行。天命赐异，源于甘星，施于阴阳，异于静动。静者天行，缓而止争；动者袍泽，速若雷霆。 捕星之计，谋策之心。甘星入世，衍造异人。奸者祸国，忠者祸身，道貌离合，莫再成臣。离经叛道，天下重分。采薇采薇，复始如周，采薇采薇，复始若秦。

讲述小魔仙夏乐彤、颜可可、白尹玥带着她们的穿梭精灵伙伴，继续守护着灵春之庭、幻夏之屿和暖秋之苑。并接纳了凛冬守护者司齐，帮助他学习知识。让司齐发现了自然的奥妙，汇聚了属于自己的蝶梦星。在这个过程中，小魔仙和司齐彼此包容，共同对抗代表了黑暗能量的夜落，建立了新的羁绊。明白了四季循环，生生不息，最终，她们凭借强大的意志，和守护四季的信念，集齐十二颗蝶梦星，用四季的自然能量，打败夜落，化解了黑暗危机。

魔气缠身的妹妹“黑莲花”夜昙公主（李兰迪饰）误入神界成为未来天妃，和缺少七情六欲的天帝长子少典有琴（陈星旭饰）从互相看不顺眼到携手拯救四界；原本命定成为天妃的姐姐青葵公主（何宣林饰）却意外嫁入魔族，与扮猪吃老虎的三皇子嘲风（陈牧驰饰）从各自心有算计，到互相理解扶持 。